RSC Tim Coulson has resigned from his role to become chief executive of the Samuel Ward Academy Trust in Suffolk.
He is the fourth RSC to resign, which means only three of the eight original RSCs are still in post.
Coulson will take over from Howard Lay to run the trust which has 15 schools.
The Samuel ward was also one of the trusts which received £200,000 from the government to become a new research hub.
Coulson said: “I was attracted to the fact that the trust is based around four towns in close geographical proximity and has those communities at its heart.
“It is also very important for individual primary, secondary and special schools to retain their identity and this is something I want to foster and enhance.”
Coulson added he was proud to have been one of the founder RSCs and “loved every minute of it”.
Nearly two thirds of Initial Teacher Training providers believe that teachers are not currently prepared to meet the government’s ambition to raise the complexity threshold for SEND pupils entering mainstream schools.
England’s councils are warning of a "ticking time bomb" in the special educational needs and disabilities (SEND) system, with new data showing deficits that could bankrupt local authorities within three years.
The regulations have been set following a second consultation and detailed collaborative working with organisations and people across deaf and hearing communities.
The Education Committee has published a letter to the Secretary of State for Education asking for more detail about the Department for Education’s work on developing its SEND reforms.